Sanctum of All

At the beginning of your upkeep, you may search your library and/or graveyard for a Shrine card and put it onto the battlefield. If you search your library this way, shuffle.
If an ability of another Shrine you control triggers while you control six or more Shrines, that ability triggers an additional time.

Island

Coral Barrier

Defender (This creature can't attack.)
When Coral Barrier enters the battlefield, create a 1/1 blue Squid creature token with islandwalk. (It can't be blocked as long as defending player controls an Island.)

Honden of Night's Reach

At the beginning of your upkeep, target opponent discards a card for each Shrine you control.

Sanctum of Stone Fangs

At the beginning of your precombat main phase, each opponent loses X life and you gain X life, where X is the number of Shrines you control.

Golgari Guildgate

Golgari Guildgate enters the battlefield tapped.
{T}: Add {B} or {G}.

Overgrown Battlement

Defender
{T}: Add {G} for each creature you control with defender.

Plains

Necropolis

Defender (This creature can't attack.)
Exile a creature card from your graveyard: Put X +0/+1 counters on Necropolis, where X is the exiled card's mana value.

Wall of Brambles

Defender (This creature can't attack.)
{G}: Regenerate Wall of Brambles.

Selesnya Guildgate

Selesnya Guildgate enters the battlefield tapped.
{T}: Add {G} or {W}.

Citadel Gate

Citadel Gate enters the battlefield tapped.
As Citadel Gate enters the battlefield, choose a color other than white.
{T}: Add {W} or one mana of the chosen color.

Steel Wall

Roving Keep

Defender
{7}: Roving Keep gets +2/+0 and gains trample until end of turn. It can attack this turn as though it didn't have defender.

Honden of Life's Web

At the beginning of your upkeep, create a 1/1 colorless Spirit creature token for each Shrine you control.

Weathered Sentinels

Defender, vigilance, reach, trample
Weathered Sentinels can attack players who attacked you during their last turn as though it didn't have defender.
Whenever Weathered Sentinels attacks, it gets +3/+3 and gains indestructible until end of turn.

Wall of Lost Thoughts

Defender
When Wall of Lost Thoughts enters the battlefield, target player mills four cards.

Mountain

Heap Gate

{T}: Add {C}.
{1}, {T}: Add one mana of any color.
{1}, {T}, Tap an untapped Gate you control: Create a Treasure token. (It's an artifact with "{T}, Sacrifice this artifact: Add one mana of any color.")

Vine Trellis

Defender (This creature can't attack.)
{T}: Add {G}.

Go-Shintai of Lost Wisdom

Flying
At the beginning of your end step, you may pay {1}. When you do, target player mills X cards, where X is the number of Shrines you control. (To mill a card, a player puts the top card of their library into their graveyard.)

Gond Gate

Gates you control enter the battlefield untapped.
{T}: Add {C}.
{T}: Add one mana of any color that a Gate you control could produce.

Glacial Wall

Wall of Bone

Defender (This creature can't attack.)
{B}: Regenerate Wall of Bone. (The next time this creature would be destroyed this turn, it isn't. Instead tap it, remove all damage from it, and remove it from combat.)

Basilisk Gate

{T}: Add {C}.
{2}, {T}: Target creature gets +X/+X until end of turn, where X is the number of Gates you control. Activate only as a sorcery.

Go-Shintai of Hidden Cruelty

Deathtouch
At the beginning of your end step, you may pay {1}. When you do, destroy target creature with toughness X or less, where X is the number of Shrines you control.

Manor Gate

Manor Gate enters the battlefield tapped.
As Manor Gate enters the battlefield, choose a color other than green.
{T}: Add {G} or one mana of the chosen color.

Sanctum of Fruitful Harvest

At the beginning of your precombat main phase, add X mana of any one color, where X is the number of Shrines you control.

Go-Shintai of Shared Purpose

Vigilance
At the beginning of your end step, you may pay {1}. If you do, create a 1/1 colorless Spirit creature token for each Shrine you control.

Wall of Hope

Defender (This creature can't attack.)
Whenever Wall of Hope is dealt damage, you gain that much life.

Wall of Resurgence

Defender
When Wall of Resurgence enters the battlefield, you may put three +1/+1 counters on target land you control. If you do, that land becomes a 0/0 Elemental creature with haste that's still a land.

Shifting Wall

Defender (This creature can't attack.)
Shifting Wall enters the battlefield with X +1/+1 counters on it.

Mirror Wall

Defender (This creature can't attack.)
{W}: Mirror Wall can attack this turn as though it didn't have defender.

Forest

Cathedral Membrane

({W/P} can be paid with either {W} or 2 life.)
Defender
When Cathedral Membrane dies during combat, it deals 6 damage to each creature it blocked this combat.

Go-Shintai of Ancient Wars

First strike
At the beginning of your end step, you may pay {1}. When you do, Go-Shintai of Ancient Wars deals X damage to target player or planeswalker, where X is the number of Shrines you control.

Wall of Air

Defender, flying (This creature can't attack, and it can block creatures with flying.)

Boros Guildgate

Boros Guildgate enters the battlefield tapped.
{T}: Add {R} or {W}.

Archers' Parapet

Defender
{1}{B}, {T}: Each opponent loses 1 life.

Mindbender Spores

Defender (This creature can't attack.)
Flying
Whenever Mindbender Spores blocks a creature, put four fungus counters on that creature. The creature gains "This creature doesn't untap during your untap step if it has a fungus counter on it" and "At the beginning of your upkeep, remove a fungus counter from this creature."

Wall of Glare

Defender (This creature can't attack.)
Wall of Glare can block any number of creatures.

Aetherflame Wall

Defender
Aetherflame Wall can block creatures with shadow as though they didn't have shadow.
{R}: Aetherflame Wall gets +1/+0 until end of turn.

Honden of Seeing Winds

At the beginning of your upkeep, draw a card for each Shrine you control.

Hover Barrier

Wall of Pine Needles

Defender (This creature can't attack.)
{G}: Regenerate Wall of Pine Needles.

Rakdos Guildgate

Rakdos Guildgate enters the battlefield tapped.
{T}: Add {B} or {R}.

Sanctum of Tranquil Light

{5}{W}: Tap target creature. This ability costs {1} less to activate for each Shrine you control.

Floriferous Vinewall

Defender
When Floriferous Vinewall enters the battlefield, look at the top six cards of your library. You may reveal a land card from among them and put it into your hand. Put the rest on the bottom of your library in a random order.

Psychic Membrane

Defender (This creature can't attack.)
Whenever Psychic Membrane blocks, you may draw a card.

Wall of Frost

Defender
Whenever Wall of Frost blocks a creature, that creature doesn't untap during its controller's next untap step.

Orzhov Guildgate

Orzhov Guildgate enters the battlefield tapped.
{T}: Add {W} or {B}.

Thunder Wall

Defender (This creature can't attack.)
Flying
{U}: Thunder Wall gets +1/+1 until end of turn.

Dazzling Ramparts

Defender
{1}{W}, {T}: Tap target creature.

Jungle Barrier

Defender (This creature can't attack.)
When Jungle Barrier enters the battlefield, draw a card.

Wall of Tears

Defender (This creature can't attack.)
Whenever Wall of Tears blocks a creature, return that creature to its owner's hand at end of combat.

Brimstone Trebuchet

Defender, reach
{T}: Brimstone Trebuchet deals 1 damage to each opponent.
Whenever a Knight enters the battlefield under your control, untap Brimstone Trebuchet.

Shimmering Barrier

Defender (This creature can't attack.)
First strike
Cycling {2} ({2}, Discard this card: Draw a card.)

Wall of Diffusion

Defender (This creature can't attack.)
Wall of Diffusion can block creatures with shadow as though Wall of Diffusion had shadow.

Wall of Omens

Defender
When Wall of Omens enters the battlefield, draw a card.

Wall of Kelp

Defender (This creature can't attack.)
{U}{U}, {T}: Create a 0/1 blue Plant Wall creature token with defender named Kelp.

Infested Roothold

Defender (This creature can't attack.)
Protection from artifacts
Whenever an opponent casts an artifact spell, you may create a 1/1 green Insect creature token.

Sanctum of Calm Waters

At the beginning of your precombat main phase, you may draw X cards, where X is the number of Shrines you control. If you do, discard a card.

Overgrown Arch

Defender
{T}: You gain 1 life.
{2}, Sacrifice Overgrown Arch: Learn. (You may reveal a Lesson card you own from outside the game and put it into your hand, or discard a card to draw a card.)

Wall of Reverence

Defender, flying
At the beginning of your end step, you may gain life equal to the power of target creature you control.

Azorius Guildgate

Azorius Guildgate enters the battlefield tapped.
{T}: Add {W} or {U}.

Portcullis Vine

Defender (This creature can't attack.)
{2}, {T}, Sacrifice a creature with defender: Draw a card.

Wall of Vines

Defender (This creature can't attack.)
Reach (This creature can block creatures with flying.)

Black Dragon Gate

Black Dragon Gate enters the battlefield tapped.
As Black Dragon Gate enters the battlefield, choose a color other than black.
{T}: Add {B} or one mana of the chosen color.

Go-Shintai of Life's Origin

{W}{U}{B}{R}{G}, {T}: Return target enchantment card from your graveyard to the battlefield.
Whenever Go-Shintai of Life's Origin or another nontoken Shrine enters the battlefield under your control, create a 1/1 colorless Shrine enchantment creature token.

Dimir Guildgate

Dimir Guildgate enters the battlefield tapped.
{T}: Add {U} or {B}.

Go-Shintai of Boundless Vigor

Trample
At the beginning of your end step, you may pay {1}. When you do, put a +1/+1 counter on target Shrine for each Shrine you control.

Wall of Distortion

Defender (This creature can't attack.)
{2}{B}, {T}: Target player discards a card. Activate only as a sorcery.

Wall of Spears

Defender (This creature can't attack.)
First strike

Wall of Blossoms

Defender
When Wall of Blossoms enters the battlefield, draw a card.

Baldur's Gate

{T}: Add {C}.
{2}, {T}: Add X mana of any one color, where X is the number of other Gates you control.

Sea Gate

Sea Gate enters the battlefield tapped.
As Sea Gate enters the battlefield, choose a color other than blue.
{T}: Add {U} or one mana of the chosen color.

Coral Colony

Defender
{1}{U}, {T}: Target player mills X cards, where X is the number of creatures you control with defender. (To mill a card, a player puts the top card of their library into their graveyard.)

Wall of Dust

Defender (This creature can't attack.)
Whenever Wall of Dust blocks a creature, that creature can't attack during its controller's next turn.

Gruul Guildgate

Gruul Guildgate enters the battlefield tapped.
{T}: Add {R} or {G}.

Fog Bank

Defender, flying
Prevent all combat damage that would be dealt to and dealt by Fog Bank.

Izzet Guildgate

Izzet Guildgate enters the battlefield tapped.
{T}: Add {U} or {R}.

Swamp

Aether Membrane

Defender; reach (This creature can block creatures with flying.)
Whenever Aether Membrane blocks a creature, return that creature to its owner's hand at end of combat.

Simic Guildgate

Simic Guildgate enters the battlefield tapped.
{T}: Add {G} or {U}.

Clockwork Drawbridge

Defender
{2}{W}, {T}: Tap target creature.

Wall of Essence

Defender (This creature can't attack.)
Whenever Wall of Essence is dealt combat damage, you gain that much life.

Gateway Plaza

Gateway Plaza enters the battlefield tapped.
When Gateway Plaza enters the battlefield, sacrifice it unless you pay {1}.
{T}: Add one mana of any color.

Sanctum of Shattered Heights

{1}, Discard a land card or Shrine card: Sanctum of Shattered Heights deals X damage to target creature or planeswalker, where X is the number of Shrines you control.

Wall of Swords

Defender (This creature can't attack.)
Flying

Wall of Denial

Defender, flying
Shroud (This creature can't be the target of spells or abilities.)

Honden of Infinite Rage

At the beginning of your upkeep, Honden of Infinite Rage deals damage to any target equal to the number of Shrines you control.

Stinging Barrier

Defender (This creature can't attack.)
{U}, {T}: Stinging Barrier deals 1 damage to any target.

Honden of Cleansing Fire

At the beginning of your upkeep, you gain 2 life for each Shrine you control.

Cliffgate

Cliffgate enters the battlefield tapped.
As Cliffgate enters the battlefield, choose a color other than red.
{T}: Add {R} or one mana of the chosen color.